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Three subspecies
P p tamilana. Southern India
P p hermosanus. Taiwan
P p tenggerensis. East Java

tamilana has recently been recognised as a separate species.
see:
NAKAE, Makoto: Taxonomic revision of five species of the family Papilionidae (Lepidoptera), Gekkan-Mushi, No. 602, Apr. 2021.

NAKAE, M. (Adam Cotton as editorial supervisor): Papilionidae of the World, Roppon-Ashi Entomological Books, Tokyo, 2021.

by the way, arjuna and hermosanus are also treated as separate species in the latter work


here the female of P. tamilana

Thanks for the info hewi. I can't keep up with all these taxonomic changes!!
Lovely female. Was never lucky enough to get hold of one.

hewi wrote: Mon Feb 20, 2023 9:35 am NAKAE, M. (Adam Cotton as editorial supervisor): Papilionidae of the World, Roppon-Ashi Entomological Books, Tokyo, 2021.

That should be "(Yasusuke Nishiyama & Adam Cotton as editorial supervisors)". Of course I cannot read the Japanese text in this book, only the English.

Adam.

hewi wrote: Mon Feb 20, 2023 9:35 am by the way, arjuna and hermosanus are also treated as separate species in the latter work

DNA analysis shows that mainland Asian Papilio paris is sister to P. karna and the Sumatran and Javan 'paris' diverged before this pair, so must be a separate species, Papilio arjuna. Similarly the southern Taiwan taxon is a separate species, P. hermosanus, whereas nakaharai from northern Taiwan belongs to P. paris.

Adam.

Here is my current listing of the Papilio paris group:

Papilio tamilana Moore, 1881 (TL: Malabar)

Papilio paris Linnaeus, 1758
ssp. paris Linnaeus, 1758 (TL: Asia [= Canton])
= majestatis Fruhstorfer, 1909 (TL: Tonkin; Annam; Tenasserim)
= splendorifer Fruhstorfer, 1909 (TL: Siam)
= f. decorosa Fruhstorfer, 1909 (TL: Sikkim)
= fa. angelicae Bryk, 1939 (TL: Assam)
= forma reductoomaculata Bryk, 1939 (TL: Assam)
ssp. chinensis Rothschild, 1895 (TL: Western China)
= f. gemmifera Fruhstorfer, 1909 (TL: Highlands of Szetchuan)
= wittmanni (Schäffler, 2004) (TL: China, Sichuan, “Rotes Becken”, Renshou)
ssp. tissaphernes Fruhstorfer, 1909 (TL: Hainan)
ssp. nakaharai Shirôzu, 1960 (TL: Showa, Taihoku Pref., N. Formosa)

Papilio hermosanus Rebel, 1906 (TL: Formosa)
= neoparis Fruhstorfer, 1908 (TL: Kagi, Formosa)
var. ♀ luli Nakahara, 1932
ab. urauri Yano, 1941
ab. divinitus Murayama, [1961]
ab. subdivinitus Murayama, [1961]
ab. superdivinitus Murayama & Shimonoya, 1965

Papilio karna C. Felder & R. Felder, 1865
ssp. discordia Nicéville, 1891 (TL: Sumatra)
= discoidia Nicéville, 1891 [Incorrect Subsequent Spelling]
= discoidea Jordan, 1909 [Incorrect Subsequent Spelling]
ssp. karna C. Felder & R. Felder, 1865 (TL: Java)
= karna C. Felder & R. Felder, 1864 [Nomen Nudum]
ssp. carnatus Rothschild, 1895 (TL: Borneo)
= karnata Fruhstorfer, 1903 [Incorrect Subsequent Spelling]
ssp. irauana Jumalon, 1975 (TL: Irauan, Puerto Princesa, Palawan)

Papilio arjuna Horsfield, 1828
ssp. battacorum Rothschild, 1908 (TL: North-East Sumatra)
ssp. dempo Okano, 1988 (TL: Mt. Dempo, S. W. Sumatra)
ssp. gedeensis Fruhstorfer, 1893 (TL: Mt. Gede, W. Java)
= prillwitzi Fruhstorfer, 1893 (Mt. Gede, W. Java)
ssp. sumbingensis Toxopeus, 1937 (TL: Soembing, ca. 1800 m)
ssp. arjuna Horsfield, 1828 (TL: Java)
= arguna Fruhstorfer, 1903 [Incorrect Subsequent Spelling]
ssp. tenggerensis Fruhstorfer, 1893 (TL: Mt. Tengger, E. Java)
= f. vern. pravieli Dufrane, 1946 (TL: Java oriental)
ssp. detanii Shimogôri, 1992 (TL: Mt. Idjen, East Jawa, Indonesia)

Adam.

hello Adam,

absolutely super, as always !!!!

did you forget the equal sign for neoparis ?

hewi wrote: Mon Feb 20, 2023 2:12 pm did you forget the equal sign for neoparis ?

Yes, I forgot to add it to my list. I do not include equal signs in my original list, but add them here for clarity. I will add it above now.

Adam.

Hello Adam,

Has P. paris tamilana been sampled in recent studies? I am curious if it has also been determined to be a separate species through mitochondrial chromosome analysis.

Ｍing.

The status of tamilana based on mt DNA sequencing has been inconclusive so far. More work is needed with additional genes.

Adam.