Interestingly Speyeria is treated as Argynnis by some websites. They are in Heliconinae subfamily so the closest would be Boloria as it is in the same Argynni tribe and then would be Heliconius and other.Chuck wrote: ↑Wed Mar 22, 2023 5:37 pm I found this, though it's 2019 and only on Speyeria. https://scholarlycommons.pacific.edu/cg ... t=uop_etds
Right up front they state they couldn't do a more comprehensive analysis due to lack of samples (specimens).
It would be interesting to know what other Nymphalid Speyeria and Argynnis are closest to; Euphydras maybe?
Euphydras is in subfamily Nymphalinae